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Arakaki, M; Christin, P A; Nyffeler, R; Lendel, A; Eggli, U; Ogburn, R M; Spriggs, E; Moore, M J; Edwards, E J (2011). Contemporaneous and recent radiations of the world's major succulent plant lineages. Proceedings of the National Academy of Sciences of the United States of America (PNAS), 108(20):8379-8384.