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Schmidt-Posthaus, H; Polkinghorne, A; Nufer, L; Schifferli, A; Zimmermann, D R; Segner, H; Steiner, P; Vaughan, L (2012). A natural freshwater origin for two chlamydial species, Candidatus Piscichlamydia salmonis and Candidatus Clavochlamydia salmonicola, causing mixed infections in wild brown trout (Salmo trutta). Environmental Microbiology, 14(8):2048-2057.