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Ashbury, Alison M; Posa, Mary Rose C; Dunkel, Lynda P; Spillmann, B; Atmoko, S Suci Utami; van Schaik, C P; van Noordwijk, M A (2015). Why do orangutans leave the trees? Terrestrial behavior among wild Bornean orangutans (Pongo pygmaeus wurmbii) at Tuanan, Central Kalimantan. American Journal of Primatology, 77(11):1216-1229.